St Laurent G, Shtokalo D, Tackett MR, Yang Z, Eremina T, Wahlestedt C, Urcuqui-Inchima S, Seilheimer B, McCaffrey TA, Kapranov P.

BMC Genomics. 2012 Sep 24;13:504. doi: 10.1186/1471-2164-13-504.

Intronic RNAs constitute the major fraction of the non-coding RNA in mammalian cells.

Argues that the intronic transcripts are long.