Chromosome Res. 2007;15(2):189-201. doi:10.1007/s10577-006-1112-z

Schulmeister A, Schmid M, Thompson EM.

Phosphorylation of the histone H3.3 variant in mitosis and meiosis of the urochordate Oikopleura dioica.

Telomeric FISH stainings with a (TTAGGG)n probe. H3.3S31 is phosphorylated during mitosis and meiosis, and not in endocycling cells. “On prometaphase chromosomes [...] H3S28P was restricted to distal chromosomal regions adjacent to telomeres whereas H3.3S31P spread throughout entire chromosomes.” “[The histone 3.3 genes] contain introns, are found outside histone clusters and their transcripts are polyadenylated.”